UNIVERSITY PARK, Pa. — Penn State knew before Saturday that replacing wide receiver Koby Howard wasn’t going to be a one-man job. Howard had emerged as the Nittany Lions’ premier explosive threat through three games, catching 12 passes for 344 yards and two touchdowns. His 28.7 yards per reception ranked among the best marks nationally,…

Penn State Needed Its Receivers to Step Up Without Koby Howard. They Didn’t.